Aspects of this topic are discussed in the following places at Britannica.
...under such diverse conditions and in such a variety of forms that their origin is uncertain. The numerous forms, with their striking variations in colour, are the product of domestication. The garden pansy (V. wittrockiana) is a hybrid, one of whose parents is V. tricolor, which is a weed of European grainfields, the other parents being V. lutea and V. altaica....
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